Boletaceae Chevall. 1826
Index Fungorum number: 80523
Type genus: Boletus L. 1753
Basidioma fleshy, boletoid, or sequestrate, occasionally phylloporoid. Pileus small to large, scaly, fibrillose, mealy, tomentose, velutinous or glabrous, dry or viscid; margin sometimes projecting; context fleshy, unchanging, bluing, browning, blackening, or reddening when bruised; hymenophore depressed, sinuate-adnexed, adnate to decurrent, lamellate or tubular whitish, pinkish, yellowish to yellow, red, or brown, unchanging or staining blue, brown, black, or red when bruised; when sequestrate, the gleba loculate, chambered or tubulose. Stipe usually fleshy, central to eccentric, cylindrical or ventricose-bulbous, solid, glabrous, or ornamented with furfuraceous to scabrous squamules, or with reticulate lines; when sequestrate, stipe often absent or surrounded by the gleba.
Basidia ballistosporic (when stipitate-pileate) or statismosporic (when sequestrate), clavate. Basidiospores roundish, broadly ellipsoid, ovate, subfusiform, or elongate subfusoid, yellowish brown, brown, or olivaceous brown to pinkish, smooth, or with different types of ornamentations. Cystidia more or less fusoid-ventricose. Hymenophoral trama boletoid or phylloporoid or intermediate. Pileipellis subcutis, ixosubcutis, trichodermium, hyphoepithelium, ixotrichodermium, ixohyphoepithelium, or composed of interwoven hyphae. Hyphae without clamp connections. Mostly ectomycorrhizal, occasionally mycoparasitic/saprotrophic